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Mugino's Meltdowner allows her to fix the electrons in the ambiguous condition that stand between a "particle" or a "wave". The manipulated electrons are then released as a white(though in this case green), sparkling ray of light that spread great destruction.
Since the high-speed particle wave is essentially electrons, a superior electrons manipulation Esper namely Misaka can manipulate it to a certain degree.
Though the opening shows it more like blocking, it maybe she's diffusing it with her ability. In hindsight, redirecting it would be more affective but it wouldn't epic enough for the opening.
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